Re: eyes in cave animals drifting away?

From: Chris Taylor (christ@ebi.ac.uk)
Date: Thu 19 Feb 2004 - 14:40:24 GMT

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    I think(...) that as a rule they go blind first, then the eyes begin to disappear as structures -- basically gross morphology is the last to go usually, so the most precise things are most vulnerable iyswim. So the metabolically expensive things don't seem to stand out -- the nerves stay attached, the mucosa open to insult. There is a gain to freeing brain for other tasks I suppose, if you have that sort of brain, as opposed to a sort of glorified ganglion.

    Keith Henson wrote:

    > At 01:21 AM 19/02/04 -0500, you wrote:
    >
    >> In a couple recent posts Keith has used the example of cave animals
    >> losing their eyes due to lack of light. In this post he tells the
    >> adaptionist tale of this loss being related to metabolic costs (ie- eyes
    >> too expensive to build so selected against):
    >>
    >> http://cfpm.org/~majordom/memetics/2000/16576.html
    >>
    >> Has he ruled out the possibility of genetic drift?
    >
    >
    > Good point.
    >
    >> In an environment
    >> that lacks light, the major selective pressure for maintaining eyes has
    >> been negated, thus mutations of eye development related genes would be
    >> selectively neutral. Populations of cave dwelling animals might be quite
    >> small.
    >
    >
    > That may be possible in some instances. In others, for insect sort of
    > things that feed on bat droppings, the population may be rather large.
    >
    >> Mutations of eye genes might accumulate, due to not being removed
    >> by selection. Eyes, as a structure, would deteriorate and the animals
    >> become blind, without metabolic cost being a significant factor in the
    >> process.
    >
    >
    > I don't know the answer to this, but do know how you would gather
    > evidence. There are a lot of different cave blind species including
    > fish and a lot of different populations as well. Has the majority shift
    > been in the direction of reduced metabolism or are there cases where the
    > eyes are just non-functional while still running the same metabolic
    > load? I.e., going blind vs the eyes shrinking to tiny dots.
    >
    > Eyes like other nerve tissue *are* expensive to operate. Additionally,
    > they are exposed and wet, making them costly in terms of protecting from
    > bacteria.
    >
    >> Futuyma's text says one possibility is that (p. 423) : "mutations that
    >> cause degeneration of an unused character become fixed by genetic drift
    >> because variations in the character are selectively neutral". Selection
    >> is another and an hypothesis is explored which supports selection but it
    >> is noted that in some instances genetic drift may play a role.
    >>
    >> Futuyma DJ. 1997. Evolutionary Biology. Sinauer Associates, Inc.
    >> Sunderland, Massachusetts
    >
    >
    > Relating this back to memetics, you would expect memetic drift where
    > there was little or no cost associated with a meme. I was considering
    > styles as an example, but that may not be drift. It is possible styles
    > don't drift but are driven in a kind of chaotic movement where the next
    > style is anything except the old one. I.e., driven to a cycle limited
    > "newness." (Women's skirts can only range from the floor to . . . . )
    > Open to speculation as to what psychological trait selected in the
    > Pleistocene could be manifesting today to drive clothing styles. Are
    > changing clothing styles a western culture feature only? Are men's
    > clothing styles more stable in different cultures?
    >
    > Keith Henson
    >
    >
    > ===============================================================
    > This was distributed via the memetics list associated with the
    > Journal of Memetics - Evolutionary Models of Information Transmission
    > For information about the journal and the list (e.g. unsubscribing)
    > see: http://www.cpm.mmu.ac.uk/jom-emit
    >
    >

    -- 
    ~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~
      Chris Taylor (christ@ebi.ac.uk)
      MIAPE Project -- psidev.sf.net
    ~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~
    ===============================================================
    This was distributed via the memetics list associated with the
    Journal of Memetics - Evolutionary Models of Information Transmission
    For information about the journal and the list (e.g. unsubscribing)
    see: http://www.cpm.mmu.ac.uk/jom-emit
    


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